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Protagonists - Saya Otonashi (main character, Chiropteran Queen) - Hagi/Haji (Saya's Chevalier) - Kai Miyagusuku (Saya's adoptive brother) - David (Saya's handler in Red Shield) Antagonists - Diva (Saya's twin sister, Chiropteran Queen) - Amshel Goldsmith (Diva's Chevalier, head of Cinq Flèches) - Solomon Goldsmith (Diva's Chevalier) - Karl Fei-Ong (Diva's Chevalier) - James Ironside (Diva's Chevalier) - Nathan Mahler (Diva's Chevalier) - Martin Bormann (Cinq Flèches associate) Red Shield - George Miyagusuku (Saya's adoptive father, Red Shield contact) - Riku Miyagusuku (Saya's adoptive brother) - Julia Silverstein (doctor) - Lewis (field operative) - Joel Goldschmidt VI (researcher, Joel Goldschmidt's descendant) - Joel Goldschmidt (original researcher) - Aston Collins (scientist) - Mao Jahana (Kai's friend, later Red Shield member) - Akihiro Okamura (reporter, aids Red Shield) - Van Argiano (scientist) - Roland Cordell (operative) - Samuel (member) - Marco (European agent) - Lina (communications specialist) - Dr. Patel (medical advisor) - Tomas (logistics coordinator) The Schiff (Artificial Chiropterans) - Moses (leader) - Karman - Lulu - Irene - Ghee - Dismas - Gestas - Darth - Gu Dr If/Gudolf Cinq Flèches & Associates - Boris (Amshel's subordinate) - Gernot (researcher) - Dietrich (scientist) - Margaret (Paris branch staff) - Annette (administrative assistant) - Hermann (security chief) - Luca (field operative) - Sophie (genetics researcher) - Sharon (Amshel's assistant) Supporting & Minor Characters - Kaori Kinjou (Riku's classmate) - Mrs. Kinjou (Kaori's mother) - Katsumi (Kai's classmate) - Yuki (Kai's baseball teammate) - Coach Tanaka (Kai's coach) - Mr. Okamura (Akihiro's father) - Sister Agnes (nun) - Captain Gardner (military officer) - Captain Torres (Philippines military officer) - Dr. Yamamoto (scientist) - Professor Tokizane (Chiropteran researcher) - Minamoto (detective)